My thinking on this, especially in view of the above ongoing air ingress is that negative pressure is NOT required, and that with any slight leak, even a weep, that air finds it far easier to gain access than for the water to seep out?. Maybe not quite alalogous to oxygen gaining access through "solid" plastic (non barrier) piping, who would have believed that but it does happen.
There was a good post on here a few years ago from someone (plumber) who had experience of two houses, one sealed, the other OV, where both had lengths of non barrier piping, the sealed one kept getting air problems but the OV one didn't, he put this down to the fact that the OV one could constantly vent any air continuously through its 22mm vent.
I think I may have a few lengths of non barrier piping in my loft area which has two rads but again never ever get air problems probably because of the OV system?.
So this means the problem could be anywhere, right?
I’ll do what I can do find the problem but short of ripping out pipe work on 3 floors I may never find it.
If the auto air vents work on the towel rail, is this equivalent to an OV system? At least air makes it’s way out of the system?